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Free radicals are generally highly reactive and participate in hydrogen abstraction, radical addition, bond scission, and annihilation reactions. For example, they can oxidize unsaturated fatty acids in cell membranes,(15-21( damage DNA, (53-57) oxidize protein amino acid side-chains,(16-19,58,59) modulate nucleotide cyclase ... .... ip.htm ... ... .... ip.htm ... ... .... ... ... ... activities,(61-67) and the action and synthesis of prostaglandins and lipoperoxides.(48,68-71). In short, radical species can attack most biological substrates from large macromolecules to smaller molecules such as catechols melanins (72-75)and the stable radical

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TABLE I. Historical perspective of biological free radicals (a few representative events)

1800's -- Vitreous chalchosis(22O) association between deafness and pigmentary abnormalities(243)

1920's -- Radical mechanisms responsible for radiation and oxygen toxicity

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1950 -- Protein antiinflammatory contaminant of hyaluronidase (Wydase) recognized by Schulte(l84)

1952 -- Application of ES R to biological materials(261)

1958-64 -- Free radical metabolites of the phenothiazines,(77,80) the melanins(73-75), Hormones(88), carbon tetrachloride(82), "Quantum Biochemistry"(134)

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